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The decade since the publication of the third edition of this volume has been an era of great progress in biology in general and the plant sciences in particular. This fourth edition reflects these developments that have transformed not only the field of biology, but also the many facets of root science. Highlights of this new edition include new understandings about roots gained in the post-genomic era, plant hormone action and signaling pathways that control root development, root-rhizosphere interactions, and modern research techniques for the field and the lab.